Biochemistry and Molecular Biophysics Graduate Group

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Graduates

Where do BMB graduates go? Many go on to postdoctoral training in top labs and then to illustrious academic or medical careers. Others play a major role in biotech and pharmaceutical companies. However, training in BMB can prepare you for many other walks of life. Below is a list of our recent Biochemistry and Molecular Biophysics graduates and their accomplishments.

2014-2015

Stephen Artim (Ph.D., August 2014)
Thesis Advisor: Mark Lemmon
Thesis Topic: Regulation and kinase activity of the Trk family of receptor tyrosine kinases
Current Position: Completing clinical rotations, University of Pennsylvania School of Veterinary Medicine
Email: sartim@vet.upenn.edu

Robert Culik (Ph.D., August 2014)
Thesis Advisor: Feng Gai
Thesis Topic: Infrared measurements of protein conformational dynamics
Postdoc: Department of Medical Genetics and Microbiology, University of Toronto, Toronto, Ontario, Canada (Lewis Kay)
Email: rculik@gmail.com

Peter Huwe (Ph.D., August 2014)
Thesis Advisor: Ravi Radhakrishnan
Thesis Topic:Uncovering the functional effects of biomolecular lmutations through computer simulations
Postdoc: Fox Chase Cancer Center (Roland Dunbrack)
Email: pjhuwe@gmail.com

David Slochower (Ph.D., August 2014)
Thesis Advisor: Paul Janmey
Thesis Topic: Multiscale simulations of phosphatidylinositol bisphosphate: understanding its biological role through physical chemistry
Postdoc: Department of Bioengineering, University of Pennsylvania, Philadelphia, PA (Paul Janmey)
Email: davids4@mail.med.upenn.edu

2013- 2014

Edward Ballister (Ph.D., May 2014)
Thesis Advisor: Michael Lampson
Thesis Topic: Inducible protein dimerization: New tools and applications to understanding the mitotic checkpoint
Postdoc: Department of Biology, University of Pennsylvania, Philadelphia, PA (Michael Lampson)
Email: ballister@gmail.com

Nicholas Bessman (Ph.D., May 2014)
Thesis Advisor: Mark Lemmon
Thesis Topic: Novel insights into the allosteric activation of the epidermal growth factor receptor
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Mark Lemmon)
Email: nichj@mail.med.upenn.edu

Ni-Ting Chiou (Ph.D., December 2013)
Thesis Advisor: Kristen Lynch
Thesis Topic: Molecular mechanisms of alternative splicing regulation: an investigation of the spliceosome repressed by HNRNP L on CD45 EXON4
Postdoc: Department of Microbiology and Immunobiology, University of California, San Francisco, CA (K. Mark Ansel)
Email: nchiou0228@gmail.com

Morgan DeSantis (Ph.D., August 2013)
Thesis Advisor: Jim Shorter
Thesis Topic: Defining the mechanism of HSP104-mediated protein disaggregation
Postdoc: Department of Cell Biology, Harvard Medical School, Boston, MA (Samara Reck-Peterson)
Email: medesantis@gmail.com

Kathryn Blakey Dupont (Ph.D., August 2013)
Thesis Advisor: Feng Gai
Thesis Topic: Spectroscopic studies of peptide-membrane interactions
Postdoc: Department of Biological Engineering, Massachusetts Institute of Technology, Cambridge, MA (Katharina Ribbeck)
Email: kathrynbdupont@gmail.com

Gabriel B. Gonzalez (Ph.D., December 2013)
Thesis Advisor: Bill DeGrado
Thesis Topic: Connectable components for protein design
Current Position: Software Engineer, Twitter, San Francisco, CA
Email: gabriel439@gmail.com

Yu-San Huoh (Ph.D., December 2013)
Thesis Advisor: Kathryn Ferguson
Thesis Topic: Investigation of substrate recognition by the Pellino E3 ubiquitin ligases
Postdoc: Program in Cellular and Molecular Medicine at Children's Hospital, Boston, MA (Sun Hur)
Email: yhuoh@hotmail.com

Tomoyasu Mani
(Ph.D., December 2013)
Thesis Advisor: Sergei Vinogradov
Thesis Topic: Magnetic field effects on molecular emissivity in solutions
Postdoc: Department of Chemistry, Brookhaven National Laboratory, Upton, NY (John R. Miller)

Melissa C. Martínez-Rivera (Ph.D. August 2013)
Thesis Advisor: Cecilia Tommos
Thesis Topic: Electrochemical characterization of TYR radicals as a function of local protein environment
Current Position: Associate Scientist, Amgen, Juncos, Puerto Rico

Nataline Meinhardt (Ph.D., May 2014)
Thesis Advisor: Doron Greenbaum
Thesis Topic: Development of specific and potent α-helical inhibitors and probes of cysteine proteases
Postdoc: Pennsylvania Drug Discovery Institute, Philadelphia, PA (Doron Greenbaum)
Email: nataline.meinhardt@gmail.com

Nisha Ninan (Ph.D., August 2013)
Thesis Advisor: Greg Van Duyne
Thesis Topic: Determinants of the SMN tudor domain interaction with the RG-tail of SmD3
Current Position: Associate Consultant, Trinity Partners, LLC, Waltham, MA
Email: nisha.s.ninan@gmail.com

Kevan Salimian (Ph.D., December 2013)
Thesis Advisor: Ben Black
Thesis Topic: Structural and mechanistic insights into centromere specification and kinetchore regulation
Current Position: Medical Student, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: kevan.salimian@gmail.com

Molly Sheehan (Ph.D., May 2014)
Thesis Advisor: P. Leslie Dutton
Thesis Topic: Engineering oxygen reactivity in heme-protein maquettes
Postdoc: to be determined
Email: sheehan.molly@gmail.com

Matthew Sochor (Ph.D., May 2014)
Thesis Advisor: Mitchell Lewis
Thesis Topic: Allostery and applications of the lac repressor
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Mitchell Lewis)
Email: msochor@mail.med.upenn.edu

Lee Solomon (Ph.D., August 2013)
Thesis Advisor: P. Leslie Dutton
Thesis Topic: The physical chemistry underlying the assembly and midpoint potential control in a series of designed protein-maquettes
Postdoc: Department of Biochemistry, University of Chicago, Chicago, IL (Tobin Sosnick)
Email: lasolomon-@uchicago.edu

Elizabeth Sweeny (Ph.D., May 2014)
Thesis Advisor: James Shorter
Thesis Topic: Structural and mechanistic insights into the yeast disaggregase Hsp104
Postdoc: Department of Biochemistry and BIophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (James Shorter)
Postdoc: Department of Biological Engineering, Massachusetts Institute of Technology, Cambridge, MA (Christopher Voigt)
Email: esweeny@gmail.com

Benjamin T. Walters (Ph.D., December 2013)
Thesis Advisor: S. Walter Englander
Thesis Topic: Large protein folding and dynamics studied by advanced hydrogen exchange methods
Postdoc: Protein Analytical Chemist II, Genentech, San Francisco, CA (Jennifer Zhang)
Email: ben@btwalters.com

2012- 2013

Joshua Allen (Ph.D., August 2012)
Thesis Advisor: Wafik El-Deiry
Thesis Topic: Efficacy and mechanistic evaluation of TIC10, a novel antitumor agent
Postdoc: Penn State Hershey Cancer Institute (Wafik El-Deiry)
Current Position: Director of Development, Oncoceutics, Inc., Bryn Mawr, PA
Email: joshjea@gmail.com

Emily Bassett (Ph.D. December 2012)
Thesis Advisor: Ben Black
Thesis Topic: Molecular mechanisms of centromere function and epigenetic propagation
Postdoc: Department of Radiation Oncology, Ohio State University, Columbus,OH (Arnab Chakravart)
Email: embassett@gmail.com

Geronda Montalvo (Ph.D. December 2012)
Thesis Advisor: William DeGrado
Thesis Topic: Design, synthesis and characterization of non-natural peptides with diverse backgrounds
Postdoc:: Research & Development, Biochemical Sciences and Engineering, Dupont, Wilmington, DE
Email: gerondam@gmail.com

Veronica Moorman (Ph.D., August 2012)
Thesis Advisor: A. Joshua Wand
Thesis Topic: Ligand-induced propagation of entropic signals across proteins
Lecturer, Department of Biology, University of Pennsylvania, Philadelphia PA, 2012 - 2014
Current Position: Assistant Professor of Biochemistry, Kettering University, Flint, MI
Email: moormanr@sas.upenn.edu

Fumin Shi (Ph.D. December 2012)
Thesis Advisor: Mark Lemmon
Thesis Topic: Signaling by non-canonical receptor tyrosine kinases ErbB3 and Ryk
Current Position: Senior Scientist, GenScript Corporation, Piscataway, NJ
Email: fumin.upenn@gmail.com

Chaim Schramm (Ph.D., August 2012)
Thesis Advisor: William DeGrado
Thesis Topic: Using structural bioinformatics to understand and design protein structures and interactions
Postdoc: Department of Biochemistry and Molecular Biophysics, Columbia University, New York, NY (Larry Shapiro/Barry Honig)

Christopher Skipwith (Ph.D. December 2012)
Thesis Advisor: X. Long Zheng
Thesis Topic: Regulation of ADAMTS13 function in hemostasis by cofactor and substrate exosite interactions
Postdoc: Department of Pharmaceutical Sciences, Northeastern University, Boston, MA (Heather Clark)
Email: c.skipwith@neu.edu

2011- 2012

Scott Bresler (Ph.D., December 2011; M.D., 2013)
Thesis Advisor: Mark Lemmon
Thesis Topic: Regulation and inhibition of the anaplastic lymphoma kinase
Current Position: Resident in Pathology, Brigham and Women's Hospital, Boston, MA
Email: sbresler@mail.med.upenn.edu

Tammer Farid (Ph.D., May 2012)
Thesis Advisor: P. Leslie Dutton
Thesis Topic: Engineering an artificial, multifunctional oxidoreductase protein maquette
Current Position: Consultant, The Boston Consulting Group, Chicago, IL
Email: tfarid.bmb@gmail.com

Jennifer Greene (Ph.D., December 2011)
Thesis Advisor: Harry Ischiropoulos
Thesis Topic: The structural proteomics of S-nitrosylation: from global identification to elucidating protein function through structural bioinformatics
Current Position: Federal government, Washington, DC
Email: jlynn.greene@gmail.com

Renee (Carrington) Martin (Ph.D. August 2011)
Thesis Advisor: Gregory Van Duyne
Thesis Topic: Structural and biochemical studies of the SMN oligomerization domain: a novel glycine zipper
Current Position: Consultant, Regulatory Affairs, Parexel International, Bethesda, MD
Email: ChezRenee05@gmail.com

David Moore (Ph.D., May 2012)
Thesis Advisor: William DeGrado
Thesis Topic: The integrin equilibrium: balancing protein-protein and protein-lipid interactions
Current Position: Medical Student, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: dtm@mail.med.upenn.edu

David Pilkinton (Ph.D., December 2011)
Thesis Advisor: Ravinder Reddy
Thesis Topic: Inhaled oxygen as a quantitative intravascular MRI contrast agent
Current Position: Medical Student, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: david.pilkinton@gmail.com

Kathryn Lorenz Sarachan (Ph.D., May 2012)
Thesis Advisor: Gregory Van Duyne
Thesis Topic:: Structural studies of the SMN-Gemin2 complex
Postdoc: National Institute of Standards and Technology (Susan Krueger)
Email: katie.sarachan@nist.gov

John Skinner (Ph.D., May 2012)
Thesis Advisors: Walter Englander and Ben Black
Thesis Topic: Protein hydrogen exchange, dynamics, and function
Postdoc: Department of Biochemistry and Molecular Biology, University of Chicago (Tobin Sosnick)
Email: skinnerj@uchicago.edu

2010 - 2011

Bryan Gibb (Ph.D. August 2010)
Thesis Advisor: Gregory Van Duyne
Thesis Topic: Requirements for catalysis in Cre recombinase
Postdoc: Department of Biochemistry and Molecular Biophysics, Columbia University, New York, NY (Eric Greene)
Email: bg2390@columbia.edu

Daniel Kulp (Ph.D. May 2010)
Thesis Advisor: William DeGrado
Thesis Topic: Using structural bioinformatics to model and design membrane proteins
Postdoc: Los Alamos National Laboratory, Los Alamos, NM (Charlie Strauss)
Current Position: Senior Research Scientist, International AIDS Vaccine Initiative, The Scripps Research Institute, La Jolla, CA
Email: dwkulp@gmail.com

Leslie Milk (Ph.D. December 2010)
Thesis Advisor: Mitchell Lewis
Thesis Topic: From structure to genetics and back: using the lac repressor to study protein-DNA interactions
Postdoc: Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, PA (JoAnne Flynn)
Current Position: Senior Project Manager, Bechtel Marine Propulsion Corporation, Pittsburgh, PA

Katarina Moravcevic (Ph.D. December 2010)
Thesis Advisor: Mark Lemmon
Thesis Topic: Investigation of phospholipid effector domains
Postdoc: Department of Neuroscience, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Amita Sehgal)
Email: katarinm@mail.med.upenn.edu

Julie Rogers (Ph.D. August 2010)
Thesis Advisor: Feng Gai
Thesis Topic: New fluorescence methods for protein folding and conformational studies
Current Position: Medical Student, Mayo Medical School, Rochester, MN
Email: juliemglass@gmail.com

Karl Schmitz (Ph.D. August 2010)
Thesis Advisor: Kathryn Ferguson
Thesis Topic: Distinct modes of binding and inhibition among anti-EGFR antibodies
Postdoc: Department of Biology, Massachusetts Institute of Technology, Cambridge, MA (Robert Sauer)
Email: schmitzk@mit.edu

Harry (Trey) W. Schroeder III (Ph.D. December 2010)
Thesis Advisors: Yale Goldman and Erika Holzbaur
Thesis Topic: Single and multiple molecule studies on the cooperation and competition of cargo-transporting motor proteins
Current Position: Resident in Surgery, Massachusetts General Hospital, Boston MA
Email: schroederh@gmail.com

J. Nathan Scott (Ph.D. August 2010)
Thesis Advisor: Jane Vanderkooi
Thesis Topic: On the nature of the hydrogen bond and biomolecular solvation
Postdoc: Department of Chemistry, Montana State University, Bozeman, MT (Patrik Callis)
Current Position: Assistant Professor of Chemistry, Saint Francis University, Loretto, PA
Email: scottjn@gmail.com

Camella Wilson (Ph.D. August 2010)
Thesis Advisor: Bryan Wolf
Thesis Topic: Characterization of the effect of Pancreatic Derived factor (PANDER) on hepatic regulation of glucose homeostasis
Postdoc: Cardiovascular Research Institute, University of California, San Francisco, CA (Ethan Weiss)
Email: camellag.wilson@gmail.com

2009 - 2010

Sung Hee Choi (Ph.D. August, 2009)
Thesis Advisor: Mark Lemmon
Thesis Topic: Understanding the regulation of the epidermal Growth Factor Receptor tyrosine kinase activity
Postdoc: Brigham and Women's Hospital, Boston, MA (Stephen Blacklow)
Email: choish2012@gmail.com

Helena Gaweska (Ph.D. December 2009)
Thesis Advisor: Dewey McCafferty
Thesis Topic: Insights into the mechanism and substrate specificity of human Lysine-Specific Demethylase 1
Postdoc: Department of Biochemistry, University of Texas Health Sciences Center, San Antonio, TX (Paul Fitzpatrick)
Email: gaweska@uthscsa.edu

John Gledhill (Ph.D. August, 2009)
Thesis Advisor: A. Joshua Wand
Thesis Topic: Optimization of multidimensional nuclear magnetic resonance spectroscopy, for resolution and sensitivity, through application of radial sampling
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia PA (A. Joshua Wand)
Current Position: Lead Scientist, Longevity Biotech, Philadelphia, PA
Email: jmgledhill@gmail.com

Sarah Chobot Hokanson (Ph.D. May 2010)
Thesis Advisor: P. Leslie Dutton
Thesis Topic: Deconvoluting the engineering and assembly instructions for Complex III activity
Postdoc: Department of Chemistry, Cornell University, Ithaca, NY (Brian Crane)
Current Position: Senior Science and Innovation Officer, British Consulate-General, Boston, MA
Email: sch95@cornell.edu

Joseph Kielec (Ph.D. December 2009)
Thesis Advisor: A. Joshua Wand
Thesis Topic: Structural studies of large integral membrane proteins in reverse micelles by solution nuclear magnetic resonance
Current Position: Life Science Specialist/Associate Consultant, L.E.K. Consulting, Boston MA
Email: jkielec@gmail.com

Joseph Laakso (Ph.D. August, 2009)
Thesis Advisor: Michael Ostap
Thesis Topic: Single molecule studies of Myosin-IB
Postdoc: National Heart, Lung & Blood Institute, National Institutes of Health, Bethesda, MD (John Hammer)
Email: joseph.laakso@gmail.com

Bruce Lichtenstein (Ph.D. May 2010)
Thesis Advisor: P. Leslie Dutton
Thesis Topic: A new approach to understanding biological control of quinone electrochemistry
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Les Dutton)
Postdoc: Department of Chemistry, University of Oxford, Oxford, United Kingdom (Ben Davis)
Email: bruce.lichtenstein@chem.ox.ac.uk

Douglas Metcalf (Ph.D. August, 2009)
Thesis Advisor: William DeGrado
Thesis Topic: Structural characterization of the integrin alphaIIb beta3 transmembrane and cytosolic domains.
Postdoc: Indiana University School of Medicine, Indianapolis, IN (Simon Conway)
Current position: Associate in the law firm of Foley Hoag LLC
Email: doug.metcalf@gmail.com

Alexei Polishchuk (Ph.D. December 2009)
Thesis Advisor: William DeGrado
Thesis Topic: Structure-based mechanism of proton transport through the influenza A/M2 protein
Current Position: Resident in Radiation Oncology, University of California, San Francisco, CA
Email: alexei.polishchuk@gmail.com

2008 - 2009

Matthew Bentley (Ph.D., August 2008)
Thesis Advisor: Dewey McCafferty
Thesis Topic: Molecular recognition and catalysis in Sortase A from Staphylococcus aureus
Postdoc: Department of Protein Engineering, Genentech, Inc., South San Francisco, CA (Andrea Cochran)
Email: matthew.bentley@gmail.com

Kelley Bethoney (Ph.D., August 2008)
Thesis Advisor: Mark Lemmon
Thesis Topic: Analysis of lipid binding domains in the dynamin family of large GPTases
Postdoc: Department of Physiology, University of Pennsylvania, Philadelphia, PA (Roberto Dominguez)
Current Position: Upper School Science Teacher, The Episcopal Academy, Newtown Square, PA
Email: kbethoney@ea1785.org

Robert Daber (Ph.D., May 2009)
Thesis Advisor: Mitchell Lewis
Thesis Topic: Rational design of novel transcriptional regulators
Postdoc: Children’s Hospital of Philadelphia Research Institute, Genetics Research Division, Philadelphia, PA (Nancy Spinner)
Current Position: Technical Director of Clinical Genomics, Center for Personalized Diagnostics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: daber@gmail.com

Chun-Chi (David) Li (Ph.D., December 2008)
Thesis Advisor: Kathryn Ferguson
Thesis Topic: Pellino proteins contain a non-canonical FHA domain that mediates the interaction with IRAK1
Postdoc: Department of Physiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Zhe Lu)
Current Position: Scientist II, Olympus Biotech, West Lebanon, NH
Email: David.Lin@olympusbiotech.com

Eric Mellon (Ph.D., December 2008)
Thesis Advisor: Ravinder Reddy
Thesis Topic: The regional cerebral metabolic rate of oxygen consumption with proton detected 17O magnetic resonance imaging during precision 17O2 inhalation in swine.
Current Position: Resident in Radiation Oncology, Moffitt Cancer Center, University of South Florida, Tampa, FL
Email: emellon@mail.med.upenn.edu

Scott Shandler (Ph.D., May 2009)
Thesis Advisor: William DeGrado
Thesis Topic: Rational design and characterization of beta-peptides, a non-natural peptidomimetic scaffold
Current Position: Founder and CEO, Longevity Biotech, Philadelphia, PA
Email: shandles@mail.med.upenn.edu

Carol Shultz (Ph.D., May 2009)
Thesis Advisor: Trevor Penning
Thesis Topic: Role of aldo-keto reductase enzymes in polycyclic aromatic hydrocarbon activation
Current Position: Associate Principal Scientist, Merck & Co., Inc., West Point, PA
Email: carol.shultz@merck.com

Walter Witschey (Ph.D., May 2009)
Thesis Advisor: Ravinder Reddy
Thesis Topic: Rotating frame magnetic resonance imaging techniques
Postdoc: University of Freiburg, Freiburg, Germany (Juergen Hennig)
Postdoc, Department of Surgery, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Robert Gorman)
Current Position: Assistant Professor of Radiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: witschey@mail.med.upenn.edu

Joy (Qingyi) Yang (Ph.D., August 2008)
Thesis Advisor: Kim Sharp
Thesis Topic: New computational tools for macromolecular simulation and modeling: charge assignment, structure modeling, and molecular dynamic simulation
Postdoc: Division of Computational and Structural Chemistry, Glaxo Smith Kline
Current Position: Scientist, Cubist Pharmaceuticals, Lexington, MA
Email: Qingyi.Yang@cubist.com

Peng Yuan (Ph.D., August 2008)
Thesis Advisor: Gregory Van Duyne
Thesis Topic: Structural land biochemical studies of site-specific recombinases
Postdoc: Laboratory of Molecular Neurobiology and Biophysics, The Rockefeller Institute, New York, NY (Roderick MacKinnon)
Email: pyuan@mail.rockefeller.edu

2007 - 2008

Gretchen Bender (Ph.D., May 2008)
Thesis Advisor: William DeGrado
Thesis Topic: De novo design and characterization of a family of metalloporphyrin proteins
Current Position: Consultant, Managed Markets, Campbell Alliance
Email: gbender_upenn@msn.com

Jamaine Davis (Ph.D., August 2007)
Thesis Advisor: Harvey Rubin
Thesis Topic: Characterization of an alternate ribonucleotide reductase small subunit, NRDF1, from Mycobacterium tuberculosis
Postdoc: National Cancer Institute at Frederick, Frederick, MD (Alex Wlodawer)
Email: jamaines@gmail.com

David Hokanson (Ph.D., December 2007)
Thesis Advisor: E. Michael Ostap
Thesis Topic: Promiscuous Myo1c and its main squeeze PIP2: it's all about the tail
Postdoc: Department of Cell and Molecular Biology, Cornell University, Ithaca, NY (Anthony Bretcher)
Email: David.Hokanson@gmail.com

David Keleti (Ph.D., May 2008)
Thesis Advisor: Mark Lemmon
Thesis Topic: Diversity in molecular recognition of phospholipids by PH domains
Current Position: Manager, Clinical Outcomes, Amerihealth Mercy Family of Companies
Email: dkeleti@gmail.com

Daryl Klein (Ph.D., May 2008)
Thesis Advisor: Mark Lemmon
Thesis Topic: Biochemical and structural studies of a secreted antagonist of the Epidermal Growth Factor Receptor
Postdoc: Harvard Medical School, Boston MA (Stephen Harrison)
Email: klein@crystal.harvard.edu

Wenkai Li (Ph.D., December 2007)
Thesis Advisor: Virginia Lee
Thesis Topic: Structural and pharmacological studies on the abnormal aggregation and normal function of tau
Current Position: Associate Principal Scientist, Pain/Migraine, Merck & Co., Inc., West Point, PA
Email: wenkai_li@merck.com

Nathaniel Nucci (Ph.D., May 2008)
Thesis Advisor: Jane Vanderkooi
Thesis Topic: On the nature of aqueous solvation in biological systems
Postdoc: Department of Biochemistry and Biophysics, University of Pennsylvania, Philadelphia, PA (A. Joshua Wand)
Current Position: Assistant Professor of Physics & Astronomy, and Translational & Biomedical Sciences, Rowan University, Glassboro, NJ
Email: nvnucci@mail.med.upenn.edu

Joanna Slusky (Ph.D., December 2007)
Thesis Advisor: William DeGrado
Thesis Topic: Sketching nature's energy function: designing helices that target TM proteins and creating software for further design
Postdoc: Department of Biochemistry and Biophysics, Stockholm University, Stockholm, Sweden (Gunnar von Heijne)
Postdoc: Fox Chase Chase Cancer Center, Philadelphia, PA (Roland L. Dunbrack, Jr.)
Email: joannas@gmail.com

Omar Zgheib (Ph.D., December 2007)
Thesis Advisor: Thanos Halazonetis
Thesis Topic: Mechanism of recruitment of p53-binding protein 1 (53BP1) to DNA double-strand breaks
Postdoc: University of Geneva, Geneva, Switzerland (Thanos Halazonetis)
Current: Graduate Program, HSBC Private Bank, Geneva, Switzerland
Email: omar.zgheib@alumni.med.upenn.edu

2006 - 2007

Jennifer Dashnau (Ph.D., May 2007)
Thesis Advisor: Jane Vanderkooi
Thesis Topic: Biological molecules as protectors against cellular damage caused by environmental extremes: the role of protectant-water interactions
Current Position: Principal Scientist, Vaccine Manufacturing Science & Commercialization, Merck & Co., Lansdale, PA
Email: jennifer.dashnau@gmail.com

Kendra Frederick (Ph.D., December 2006)
Thesis Advisor: A. Joshua Wand
Thesis Topic: Protein dynamics and thermodynamics: a view emerging from NMR relaxation studies on the calmodulin system
Postdoc: Whitehead Institute, Cambridge (Susan Lindquist)
Email: kendrakf@wi.mit.edu

Soo Hwa Kim (Ph.D., December 2006)
Thesis Advisor: Marcos Milla
Thesis Topic: A functional study of the cytoplasmic domain of tumor necrosis factor-α converting enzyme
Current Position: Patent Agent, Ropes & Gray, LLP
Email: soohwakim@alumni.upenn.edu

Benjamin North (Ph.D., December 2006)
Thesis Advisor: William DeGrado
Thesis Topic: Application of the Crick parameterization to de novo protein design
Postdoc: Institute for Cancer Research, Fox Chase Cancer Center, Philadelphia, PA (Roland Dunbrack, Jr.)
Email: bnorth@mail.med.upenn.edu

Lumelle Schneeweis (Ph.D., May 2007)
Thesis Advisor: Paul Axelsen
Thesis Topic: Structural and functional analysis of osteoprotegerin and apolipoprotein E
Current Position: Research Investigator, Bristol-Myers Squibb, Princeton, NJ
Email: lumelle.schneeweis@bms.com

Rachel Signarvic (Ph.D., December 2006)
Thesis Advisor: William DeGrado
Thesis Topic: The de novo design of molecular switches: manipulation of secondary structure to modulate conformational stability
Postdoc: Department of Medicine, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Skip Brass)
Email: signarvi@mail.med.upenn.edu

2005 - 2006

Andrew Avarbock (Ph.D., December 2005)
Thesis Advisor: Harvey Rubin
Thesis Topic: Functional regulation of RelMtb: the stringent response protein responsible for initiating dormancy in Mycobacterium tuberculosis
Current Position: Resident, Department of Dermatology, New York Presbyterian-Columbia University, New York, NY

Sabrina Bedard (Ph.D., May 2006)
Thesis Advisor: S. Walter Englander
Thesis Topic: Hydrogen exchange studies of staphylococcal nuclease folding and unfolding
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Josh Wand)
Past Position, Research Specialist, Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Current Position: Investigator, Research and Development, Biophysics), GlaxoSmithKline, Collegeville, PA
Email: sabrina.bedard@gmail.com

Mary Fitzgerald (Ph.D., December 2005)
Thesis Advisor: Ronen Marmorstein
Thesis Topic: Everted half-site recognition by Leu3: structure/function relationships of a Zn2Cys6 binuclear cluster motif
Postdoc: Rutgers University Center For Advanced Biotechnology And Medicine, Busch Campus, Piscataway, NJ
Email: mfitzger@cabm.rutgers.edu

Brenda Frankel (Ph.D., August 2005)
Thesis Advisor: Dewey G. McCafferty
Thesis Title: Inhibition and catalytic mechanism of the Staphylococcus aureus
Postdoc: Albert Einstein College of Medicine (John Blanchard)
Current Position: Senior Research Scientist, OSI Pharmaceuticals (Astellas Pharmacology), Melville, NY
Email: bpetrone@aecom.yu.edu

G. Scott Pesiridis (Ph.D., May 2006)
Thesis Advisor: Gregory Van Duyne
Thesis Topic: Allosteric regulation of PRMT5 by cooperative interactions of the plCln-SmD3 complex
Postdoc: Department of Pathology & Laboratory Medicine, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Virginia Lee)
Current Position: Investigator, Molecular Discovery Research, GlaxoSmithKline, Collegeville, PA
Email: g.s.pesiridis@gsk.com

2004 - 2005

Ari Goldberg (Ph.D., December 2004)
Thesis Advisor: John Leigh
Thesis Topic: New approaches to higher resolution images in Diffusion Optical Tomography
Resident, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Current Position: Assistant Professor of Radiology, Penn State Geisinger Health System, Wilkes Barre, PAEmail: arig@mail.med.upenn.edu

Patricia E. Gonzales (Ph.D., May 2005)
Thesis Advisor: Marcos Milla
Thesis Topic: Inhibition of the tumor necrosis factor-alpha converting exzyme by its pro domain
Current Position: Research Assistant, Pontificia Univeresidad Catolica del Peru, Lima, Peru
Email: pgonzales@pucp.edu.pe

Justin Hulvershorn (Ph.D., December 2004)
Thesis Advisor: John Leigh
Thesis Topic: Imaging neural activity: improved localization, quantitation, and temporal resolution using novel fMRI contrasts
Current Position: Chief Scientific Officer, Mirador Biomedical, Seattle, WA
Email : jhulvershorn@miradorbiomedical.com

Megan King (Ph.D., December 2004)
Thesis Advisor: Mark Lemmon
Thesis Topic: The role of targeting and effector domains in the functions of dynamin-family GTPases
Postdoc: The Rockefeller University, New York, NY (Guenter Blobel)
Current Position: Assistant Professor, Department of Cell Biology, Yale University, New Haven, CN
Email: megan.king@yale.edu

Chia-Shang (Jason) Liu (Ph.D., August 2005)
Thesis Advisor: Mark Elliott
Title: The nature of the blood oxygen level dependent signal in functioning magnetic resonance imaging: influences and intrinsic functional information
Current Position: Radiology Resident, Albert Einstein Medical Center, Philadelphia, PA

Yingli Ma (Ph.D., August 2005)
Thesis Advisor: Gregory D. Van Duyne
Thesis Title: Structural and biochemical studies of the Gemin6/7 heterodimer from the human SMN complex
Postdoc: The Howard Hughes Medical Institute, The Rockefeller University, New York, NY (Gunter Blobel)
Current Position: Senior Scientist, Research & Development, GlaxoSmithKline (Bioinformatics and Structural Biology), China
Email: yingli.ma@gmail.com

Mohita Mohan (Ph.D., August 2005)
Thesis Advisor: Marcos Milla
Thesis Title: Substrate specificity is a determinant of ectodomain shedding by Tumor Necrosis Factor Alpha Converting Enzyme (TACE)
Current Position: Market Planning Manager, Genentech, Inc. (Biotechnology), South San Francisco, CA
Email: mohitamohan@yahoo.com

Heather Pinkett (Ph.D., December 2004)
Thesis Advisor: Mitchell Lewis
Thesis Topic: Structural determination of the phage 186 repressor protein
Postdoc: California Institute of Technology, Pasadena, CA (Douglas Rees)
Current Position: Assistant Professor, Department of Biochemistry, Molecular Biology, and Cell Biology Northwestern University, Chicago, IL
Email: h-pinkett@northwestern.edu

Avram Slovic (Ph.D., December 2004)
Thesis Advisor: William DeGrado
Topic: Water-solubilization of integral membrane proteins
Postdoc: Columbia University, New York, NY (James Rothman)
Current Position: Biofuels Program Manager, Amyris Biotechnologies, Sao Paulo, Brazil
Email: slovic@amyris.com

Lawrence Szewczuk (Ph.D., December 2004)
Thesis Advisor: Trevor Penning
Thesis Topic: Selective mechanism-based inactivation of COX-1 by m-hydroquinones
Postdoc: Johns Hopkins University, Baltimore, MD (Philip Cole)
Current Position: Principal Scientist, GlaxoSmithKliine (High-Throughput Screening), Collegeville, PA
Email: suzak_44@yahoo.com

Jason Wan (Ph.D., May 2005)
Thesis Advisor: Joseph Jarrett
Thesis Topic: Flavodoxin-dependent biotin synthase
Postdoc: NIDDK, National Institutes of Health (Jeremy Berg)
Current Position, Program Specialist, Integrative Biology and Infectious Diseases Branch, Division of Extramural Research, National Institute of Dental and Craniofacial Research, National Institutes of Health, Bethesda, MD
Email: JasonWan@nidcr.nih.gov

2003 - 2004

Mitchell Berger (Ph.D., May 2004)
Thesis Advisor: Mark Lemmon
Thesis Topic: Mechanisms of ErbB receptor oligomerization
Current Position: Clinical Lecturer, Female Pelvic Medicine and Reconstructive Surgery, Division of Gynecology, Department of Obstetrics and Gynecology, University of Michigan, Ann Arbor, MI
Email: mitcherg@med.umich.edu

P. Dayanand Borge (Ph.D., December 2003)
Thesis Advisor: Bryan Wolf
Thesis Topic: IRS-1 inhibition of SERCA: a potential molecular interaction mechanism for regulation of insulin secretion and calcium homeostasis in pancreatic β-cells
Postdoc: Department of Pediatrics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Bryan Wolf)
Current Position: Clinical Fellow, Department of Transfusion Medicine, NIH Clinical Center, Bethesda MD
Email: dayandborge@yahoo.com

Dawn Eastmond (Ph.D., May 2004)
Thesis Advisor: Stephen Liebhaber
Thesis Topic: Alteration of gene expression by specially designed RNA decoys
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Hillary Nelson)
Current Position: Director of Education, Office of Graduate Studies, Scripps Research Institute Kellogg School of Science and Technology La Jolla, CA

Jennifer Gronlund Haluko (Ph.D., December 2004)
Thesis Advisor: Dewey McCafferty
Thesis Topic:Studies on the mechanism of histone deacetylase enzymes
Present Position: Research Scientist, Colgate-Palmolive Company, Piscataway, NJ (Malcolm Williams)
Email: jennikins@gmail.com

Vladimira Heredia (Ph.D., December 2004)
Thesis Advisor: Trevor Penning
Thesis Topic: Determinants of the rate-limiting events in 3α-hydroxysteroid dehydrogenase (3α-HSD) catalysis
Postdoc: Department of Biochemistry, Pfizer, Inc., San Diego, CA (Shaoxian Sun)
Current Position: Assistant Professor, Department of Chemistry, Temple University, Philadelphia, PA
Email: tue40781@temple.edu

Li Jia (Ph.D., May 2004)
Thesis Advisor: Jonathan Raper
Thesis Topic: Slits are chemorepellents for retinal ganglion cell axons and trunk neural crest cells
Postdoc: NIDDK, NIH, Bethesda, MD (Douglas Forrest)
Past Position: Postdoctoral Fellow, National Eye Institute, NIH, Porter Neuroscience Research Center, Bethesda, MD (Wei Li)
Current Position: Research Fellow, Department of Retinal Neurophysiology, National Eye Institute, NIH, Bethesday, MD
Email lijia@yahoo.com

2002 - 2003

Linh Hoang (Ph.D., May 2003)
Thesis Advisor: Walter Englander
Thesis Topic: Transient intermediate units in protein folding
Current Position: MD Director, Personalized Medicine, Life Technologies

Shihe (Shawn) Hou (Ph.D., December 2002)
Thesis Advisor: Robert Ricciardi
Thesis Topic: Transcriptional regulation of MHC class I genes in adenovirus transformed cells: COUP-TF and NF-KB. A tale of two transcription factors
Postdoc: Department of Microbiology, University of Pennsylvania School of Dental Medicine, Philadelphia, PA (Robert Ricciardi)
Current Position: Scientist, Abraxis Bioscience, Inc., Los Angeles, CA
Email: shou@cpcus.jnj.com

Wai Lau (Ph.D., May 2003)
Thesis Advisor: William DeGrado
Thesis Topic: Folding dynamics and membrane fusion mediated by model coiled-coil peptides
Postdoc: Massachusetts Institute of Technology, Cambridge, MA (K. Dane Wittrup)
Current Position: Scientist, Adnexus Therapeutics, Waltham, MA
Email: waileunglau@gmail.com

Anthony Lee (Ph.D., May 2003)
Thesis Advisor: Mark Lemmon
Thesis Topic: Study of the dynamin PH domain and its role in receptor-mediated endocytosis
Postdoc: University of California, Los Angeles, CA (Linda Liau)
Current Position: Law Student, University of Wisconsin, Madison, WI

Wei Li (Ph.D., August 2002)
Thesis Advisor: Rebecca Taub
Thesis Topic: The role of interleukin-6 and STAT3 in liver regeneration
Postdoc: Genomics Institute of the Novartis Research Foundation (GNF) & Department of Cell Biology, The Scripps Research Institute, La Jolla, CA (Claudio Joazeiro)
Current Position: Scientist, Rigel Pharmaceuticals Inc., South San Francisco, CA

Omar Manuar (Ph.D., December 2002)
Thesis Advisor: Dwight Jaggard
Thesis Topic: The effect of parametric direction on scattering from wire loops
Postdoc: University of Pennsylvania, Department of Bioengineering, Philadelphia, PA (Dwight Jaggard)
Current Position: Unknown

Francesco Vanzi (Ph.D., December 2002)
Thesis Advisor: Yale Goldman
Thesis Topic: Mechanical studies of single ribosome/mRNA complexes
Current Position: Tenure Researcher, Department of Biology, University of Florence, Italy
Email: fvanzi@lens.unifi.it

Xun Wang (Ph.D., May 2003)
Thesis Advisor: Mortimer Poncz
Thesis Topic: Control of megakaryocyte-specific gene expression by GATA-1 and FOG-1: role of Ets transcription factors
Postdoc: University of Pennsylvania Department of Pediatrics, Philadelphia, PA (Mortimer Poncz)
Current Position: Sr. Research Scientist, Quest Pharmaceutical Services, Newark, DE

Jeongsik Yong (Ph.D., August 2002)
Thesis Advisor: Gideon Dreyfuss
Thesis Topic: Diverse functions of RNA-binding proteins in the biogenesis of ribonucleoprotein particles
Postdoc: Howard Hughes Medical Institute, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Gideon Dreyfuss)
Current Position: Assistant Professor of Biochemistry, Molecular Biology and Biophysics, University of Minneapolis College of Biological Sciences, Twin Cities, Minneapolis, MN
Email: jeongsik@mail.med.upenn.edu

2001 - 2002

Uma Duvvuri (Ph.D., May 2002)
Thesis Advisor: John Leigh
Thesis Topic: Magnetic resonance relaxation studies of osteoarthritis and tumor response to chemotherapy
Residency: University of Pittsburgh Medical School, Pittsburgh, PA
Fellowships: University of Texas MD Anderson Cancer Center, Houston, TX
Current Position: Assistant Professor, Department of Otolaryngology, University of Pittsburgh Medical Center, Pittsburgh, PA
Email: duvvuriu@upmc.edu

Maria Fernandez-Seara (Ph.D., May 2003)
Thesis Advisor: Felix Wehrli
Thesis Topic: Structural and functional implications of bone water and mineral density studied by NMR
Postdoc: Functional Imaging Laboratory, London, UK (Robert Turner)
Current Position: Staff Scientist, Center for Applied Medical Research (CIMA), University of Navarra, Pamplona, Spain
Email: mfseara@unav.es

Kelly Gallagher (Ph.D., May 2002)
Thesis Advisor: Kim Sharp
Thesis Topic: Electrostatic contributions to heat capacity changes and an analysis of thermal hysteresis protein hydration using the random network model
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Kim Sharp)
Current Position: Assistant Professor,, Department of Chemistry & Biochemistry, SUNY College at Oneonta, Oneonta, NY
Email: gallagkr@oneonta.edu

Steve Huang (Ph.D., December 2001)
Thesis Advisor: P. Leslie Dutton
Thesis Topic: Structure of a molecular maquette scaffold
Residency: Nuclear Medicine, Hospital of The University of Pennsylvania, Philadelphia, PA
Current Position: Physician, Cleveland Clinic Imaging Institute, Cleveland, OH

Lisa Lai (Ph.D., December 2001)
Thesis Advisor: Sandra Holloway
Thesis Topic: Characterization of CDC16-183: a novel APC mutant which activates the spindle checkpoint
Postdoc: Department of Pathology, University of Washington, Seattle, WA (Peter Ravinovich)
Current Position: Research Scientist, University of Washington, Seattle, WA
Email: lail@u.washington.edu

Shuchun (Donald) Li (Ph.D., December 2001)
Thesis Advisor: John Leigh
Thesis Topic: New methods in MRI susceptometry: theory, experiments, application
Postdoc: Research Associate, Department of Physics, University of Pennsylvania, Philadelphia, PA
Current Position: Unknown

Jianping (Jenny) Ni (Ph.D., December 2001)
Thesis Advisor: Gregory Van Duyne
Thesis Topic: Structural studies of arginine repressor from Bacillus stearothermophilus
Postdoc: Department of Biochemistry and Biophysics, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (Mitchell Lewis)
Current Position: Unknown

Doris Ohliger (Ph.D., August 2001)
Thesis Advisor: Robert Forster
Thesis Topic: Carbonic anhydrase: how does this zinc enzyme work? Carbon-13 magnetization transfer studies of carbon dioxide-bicarbonate exchange
Current Position: HP Technician, Safety Office, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: doris@ehrs.upenn.edu

Nisha Palackal (Ph.D., May 2002)
Thesis Advisor: Trevor Penning
Thesis Topic: Role of human aldo-keto reductases in PAH activation
Past Position: Research Scientist, Cayman Chemical Co., Ann Arbor, MI
Current Position: Group Leader, Hospira, Inc., Lake Forest, IL

Steven Stayrook (Ph.D., May 2001)
Thesis Advisor: Mitchell Lewis
Thesis Topic: Crystal structure of human sterol carrier protein 2
Postdoc: Howard Hughes Medical Institute; University of Pennsylvania, Philadelphia, PA
Current Position: Senior Research Investigator; Department of Biochemistry and Biophysics; Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: stayrook@mail.med.upenn.edu

Christopher Summa (Ph.D., May 2002)
Thesis Advisor: William DeGrado
Thesis Topic: Computational methods and their applications for de novo functional protein design and membrane protein solubilization
Postdoc: Department of Structural Biology, Stanford University School of Medicine, Stanford, CA (Michael Levitt)
Current: Assistant Professor of Computer Science, University of New Orleans. LA and Assistant Professor, Research Institute for Children, New Orleans, LA
Email: csumma@cs.uno.edu

2000 - 2001

Marcus Bell (Ph.D., December 2000)
Thesis Advisor: Yale Goldman
Thesis Topic: Polarized fluorescence depletion reports orientation distribution and rotational dynamics of muscle cross-bridges
Postdoc: Department of Pathology, Thomas Jefferson University, Philadelphia, PA (Robert Barsotti)
Current Position: Associate Professor of Biomedical Sciences, Philadelphia College of Osteopathic Medicine, Philadelphia, PA
Email: MarcusBe@pcom.edu

Sridhar Charagundla (Ph.D., December 2000)
Thesis Advisor: John S. Leigh
Deceased

David Clayton (Ph.D., December 2000)
Thesis Advisor: John Leigh
Thesis Topic: Multinuclear magnetic resonance techniques for the study of brain metabolism in vivo
Postdoc: Department of Radiology, Stanford University School of Medicine, Lucas Center for MR Imaging and Spectroscopy, Stanford, CA (Dan Spielman)
Current Position: Physical Science Research Associate, Stanford University, Stanford, CA
Email: dclayton@stanford.edu

Adrienne Clements (Ph.D., May 2001)
Thesis Advisor: Ronen Marmorstein
Thesis Topic: Biophysical characterization and structural studies of two E1A-interacting proteins: the pRB tumor suppressor and the PCAF coactivator
Postdoc: Wistar Institute, Philadelphia, PA 19104 (Ronen Marmorstein)
Current Position: Associate Scientific Director, Janssen Research and Development, Johnson and Johnson , Springhouse, PA
Email: adrienne.clements.egan@verizon.net

Masaru Ishii (Ph.D., August 2001)
Thesis Advisor: John Leigh
Thesis Topic: Photon diffusion imaging in model and biological systems
Current Position: Assistant Professor, Department of Otolaryngology–Head and Neck Surgery, Johns Hopkins University, Baltimore, MD

Kanli Jiang (Ph.D., August 2001)
Thesis Advisor: John Wolfe
Thesis Topic: Genetic engineering of a therapeutic lysosomal enzyme to improve its delivery into the central nervous system
Current Position: Physician, Pennsylvania Hospital, Philadelphia, PA

Jonathan Kaufman (Ph.D., December 2000)
Thesis Advisor: John S. Leigh
Thesis Topic: Magnetic resonance studies of cartilage osmotic and mechanical properties
Postdoc: Department of Radiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA (John S. Leigh)
Current Position: CEO, Lipella Pharmaceuticals, Pittsburgh, PA
Email: jhk@lipella.com

Raymond Trievel (Ph.D., December 2000)
Thesis Advisor: Ronen Marmorstein
Thesis Topic: Crystal structure, catalytic mechanism, and substrate specificity of the GCN5 histone acetyltransferase family
Postdoc: National Institutes of Health, Bethesda, MD (James Hurley)
Current Position: Associate Professor of Biological Chemistry, University of Michigan, Ann Arbor, MI
Email: rtrievel@umich.edu

Ravi Venkataramani (Ph.D., May 2001)
Thesis Advisor: Ronen Marmorstein
Thesis Topic: Crystal structure and mutational analysis of cell cycle regulator, p18INK4c
Postdoc: The Wistar Institute, Philadelphia, PA (Ronen Marmorstein)
Current Position: Associate Director, ImClome Systems, Inc.

Graduates before December 2000 – thesis advisor and current position

Ari Borthakur (Ph.D., August 2000) Thesis Advisor: Ravinder Reddy
Senior Research Investigator, Department of Radiology, Perelman School of Medicine at the University of Pennsylvania
& MBA candidate, The Wharton School, University of Pennsylvania, Philadelphia, PA
Email: borthaku@mail.med.upenn.edu

Xiaoxi Chen (Ph.D., December 1999) Thesis Advisor: P. Leslie Dutton
Research Scientist, Phylos, Inc., Watertown, MA
Email: kchen@phylos.com

Ivan Dimitrov (Ph.D., May 2000) Thesis Advisor: John Leigh
Senior Scientist, Phillips Medical Systems, Dallas, TX
Email: ivan.dimitrov@utsw.edu

Mark Elliott (Ph.D., May 2000) Thesis Advisor: John Leigh
Senior Staff Investigator, Department of Radiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: mark@mail.mmrrcc.upenn.edu

Tara Leigh Beers Gibson (Ph.D., December 1999) Thesis Advisor: Pinchas Cohen
Director, Medical Communications at Scientific Connexions, Newtown, PA

Scott Keven Gibson (Ph.D., December 1999)
Thesis Advisor: Paul Liebman
Deceased

Robert Greenman (Ph.D., May 2000) Thesis Advisor: Robert Lenkinski
Staff Physicist, Beth Israel Deaconess Medical Center and Instructor, Harvard Medical School, Boston, MA
Email: rgreenma@caregroup.harvard.edu

Anne Marie Grosset (Ph.D., May 2000) Thesis Advisor: P. Leslie Dutton
Head, Polymer Chemistry, SICPA, Basel, Switzerland

Feng Guo (Ph.D., May 2000) Thesis Advisor: Gregory Van Duyne
Associate Professor of Biological Chemistry, University of California, Los Angeles, Los Angeles, CA
Email: fguo@mbi.ucla.edu

Alan Haas (Ph.D., August 2000) Thesis Advisor: P. Leslie Dutton
A.L.M. in Psychology degree candidate, Harvard Extension School, Boston, MA
Email: ahaas@hsph.harvard.edu

Elizabeth Hopper–Borge (Ph.D., May 1999) Thesis Advisor: Tomoko Ohnishi
Assistant Professor (Developmental Therapeutics), Fox Chase Cancer Center, Philadelphia, PA
Email: Elizabeth.Hopper@fccc.edu

Astrid Kral (Ph.D., May 2000) Thesis Advisor: Charles Omer
Senior Research Biochemist, Merck Research Labs, West Point, PA
Email: astrid_kral@merck.com

Lin Li (Ph.D., May 2000) Thesis Advisor: John Leigh
Research Assistant Professor of Radiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA
Email: linli@mail.med.upenn.edu

Xinmin Li (Ph.D., August 2001) Thesis Advisor: Ronen Marmorstein
Research Associate, University of Pittsburgh, Pittsburgh, PA (Structural Biology)
Email: xil85@pitt.edu

Daniel Scolnick (Ph.D., May 2000) Thesis Advisor: Thanos Halazonetis
Patent Attorney, Pepper Hamilton LL, Philadelphia, PA.
Email: scolnickd@pepperlaw.com

Jeffrey Souris (Ph.D., December 1999) Thesis Advisor: John Leigh
Research Assistant Professor of Radiology and Technical Director, Optical Imaging Core, Biological Science Division, The University of Chicago, Chicago, IL
Email: sourisj@uchicago.edu

Change Tan (Ph.D., August 2000) Thesis Advisor: Peter Klein
Assistant Professor of Biological Sciences, University of Missouri-Columbia, Columbia, MO
Email: tanc@missouri.edu

Zhenyu Tang (Ph.D., May 2000) Thesis Advisor: Thomas Kadesch
Managing Partner, Minerva Capital Partners Ltd., Beijing, China

Scott T. R. Walsh (Ph.D., August 2000) Thesis Advisor: William DeGrado
Assistant Professor, Department of Cell Biology & Molecular Genetics, University of Maryland, College Park, MD
Email: swalsh12@umd.edu

Jie Yao (Ph.D., December 1999) Thesis Advisor: Daniel Kessler
Email: jj_yao@yahoo.com

Zhenyu Tang (Ph.D., May 2000) Thesis Advisor: Thomas Kadesch
Graduated from Kellogg School of Management

Nabil Chehab (Ph.D., December 1999) Thesis Advisor: Thanos Halazonetis
Principal Scientist, Imclone Systems, Eli Lilly and Company
Email: chehabn@mskcc.org

Cheryl Clarkin (Ph.D., August 1999)
Thesis Advisor: Barbara Weber
Physician, Department of Pediatrics, Bryn Mawr Hospital, Bryn Mawr, PA

Farhad Kosari (Ph.D., May 1999) Thesis Advisor: Thomas Kleyman
Assistant Professor of Biochemistry and Molecular Biology, Cancer Center at Mayo Clinic, Rochester, MN
Email: Kosari.Farhad@mayo.edu

Kai Lin (Ph.D., August 1999) Thesis Advisor: Robert Ricciardi
Director of Biology, Permeon Biologics, Cambridge, MA
Email: kai.lin@novartis.com

Lin Liu (Ph.D., May 1999) Thesis Advisor: Shelley Berger

Karyn O'Neil (Ph.D., December 1998) Thesis Advisor: William DeGrado
CSO, Centyrex Venture, Johnson & Johnson
Email: koneil@cntus.jnj.com

Naoyuki Saito (Ph.D., December 1998)
Thesis Advisor: Yvonne Paterson
Assistant Professor, Departments of Radiation Medicine and Pharmacology & Therapeutics, Roswell Park Cancer Institute and Assistant Professor, SUNY, Buffalo, NY
Email: ngsaito@buffalo.edu

Julia Shifman (Ph.D., December 1998) Thesis Advisor: P. Leslie Dutton
Principal Investigator, The Alexander Siberman Institute of Life Sciences, The Hebrew University of Jerusalem, Jerusalem, Israel
Email: jshifman@cc.huji.ac.il

Liat Shimoni (Ph.D., December 1998) Thesis Advisor: Jenny Glusker
Clinical Data Analyst, University of Pennsylvania Health System, Philadelphia, PA
Email: Liat.Shimoni@uphs.upenn.edu

Mark Shtilerman (Ph.D., August 1999) Thesis Advisor: Walter Englander
Associate in Pharmaceutical and Biotech Patent Litigation, Paul Hastings
Email: mshtilerman@dl.com

Jeffrey Taylor (Ph.D., December 1998) Thesis Advisor: Roger Burnett

Alissa Woska (Ph.D., December 1998) Thesis Advisor: Michael. Klein
Research Chemist, Hercules, Inc., Wilmington, DE

Hui Zhang (Ph.D., May 1999) Thesis Advisor: Roland G. Kallen
Associate Professor, Department of Pathology, Johns Hopkins University, Baltimore, MD
Email: hzhang32@jhmi.edu

Jinsong Zhang (Ph.D., August 1999) Thesis Advisor: Mitchell Lazar
Assistant Professor of Cancer and Cell Biology, Vontz Center for Molecular Studies, U. of Cincinnati Medical Center, Cincinnati, OH
Email: zhangj5@ucmail.uc.edu

Enn-Ling Chen (Ph.D., August 1998) Thesis Advisor: John Leigh
Assistant Research Professor, Division of Cardiology, Department of Medicine, Duke University Medical Center, Durham, NC

Douglas Fletcher (Ph.D., May 1998) Thesis Advisor: John Haselgrove
Lieutenant, United States Navy and Assistant Professor of Radiology, Uniformed Services University of the Health Sciences, Bethesda, MD
Email: dfletcher@usuhs.mil

Joseph Jez (Ph.D., May 1998) Thesis Advisor: Trevor Penning
Associate Professor, Department of Biology, Washington University, St. Louis, MO
Email: jjez@wustl.edu

Stephen Mulholland (Ph.D., December 1997) Thesis Advisor: P. Leslie Dutton
Research Scientist, Department of Chemistry and Biochemistry, Biacore, AB, Cambridge, MA
Email: semulholland@earthlink.net

J. Michael Sauder (Ph.D., May 1998) Thesis Advisor: Heinrich Roder
Senior Scientist, Eli Lilly, San Diego CA
Email: sauder_j_michael@lilly.com

Sean-Patrick Scott (Ph.D., August 1998) Thesis Advisor: Jacqueline Tanaka
Professor Investigator, Escuela de Medicina, Instituto Tecnologico y de Estudios
Superiores de Monterrey, Monterrey, Nuevo León, México
Email: spscott@itesm.mx

Mark Tang (Ph.D., August 1997) Thesis Advisor: Trevor Penning
Operations Manager, M+W Zander, Ltd., Bangkok, Thailand
Email: mswtang@yahoo.com

Robert Tjin Tham Sjin (Ph.D., August 1998) Thesis Advisors: Barbara Hoffman and Dan Liebermann
Scientist, Avila Therapeutics, Waltham, MA
Email: r_tjin@hotmail.com

Lian Wang (Ph.D., August 1997) Thesis Advisor: Shelley Berger
Principal Scientist, Department of Biology, Boehringer Ingelheim Pharmaceuticals, Inc., Ridgefield, CT
Email: lwang@rdg.boehringer-ingelheim.com

Seth Hopkins (Ph.D., May 1997) Thesis Advisor: Yale Goldman
Research Director, Preclinical Research and Translational Medicine, Sunovion Pharmaceuticals, Marlborough, MA
Email: seth.hopkins@sunovion.com

Zhiqiang Liu (Ph.D., August 1997) Thesis Advisor: Truman Brown
Consultant, CB Technologies, Inc, Malvern, PA

Glenn Walter (Ph.D., May 1997) Thesis Advisor: John Leigh
Associate Professor of Physiology and Functional Genomics, University of Florida, Gainesville, FL
Email: glennw@ufl.edu

Young-Han Song (Ph.D., May 1998) Thesis Advisor: Nancy Cooke
Associate Professor, Ilsong Institute of Life Science, Hallym University, Gangwon-do, Korea
Email: ysong@hallym.ac.kr

Alan Stein (Ph.D., August 1998) Thesis Advisor: John Leigh
Vice President of Healthcare Technology, Autonomy Corp (An HP company), Cherry Hill, NJ
Email: alan@medcosm.com