University of Wales, Biochemistry, Ph.D., 1967
2008 Member MRSEC Committee (NSF): Assembly of Nanoscale Protein Materials
1994- 2008 Chairman, Department of Biochemistry and Biophysics, University of Pennsylvania
1993-1994 Acting Chairman, Department of Biochemistry and Biophysics, University of Pennsylvania
1991- Director, Johnson Research Foundation, University of Pennsylvania
1981- Professor, Department of Biochemistry and Biophysics, University of Pennsylvania
1976 Associate Professor, Department of Biochemistry and Biophysics, University of Pennsylvania
1971 Assistant Professor, Johnson Research Foundation, University of Pennsylvania
1968 Post Doctoral Fellow with Britton Chance, Johnson Research Foundation, University of Pennsylvania
1967 Post Doctoral Fellow with W. Charles Evans, Department of Biochemistry and Soil Science, University of Wales
2001- Visiting Senior Fellow, St. John’s College, Oxford
1995-1998 Visiting Professor, University College, London
1994- Visiting Professor, Imperial College, London
1975-1978 Visiting Scientist, Bell Research Laboratories, Murray Hill, New Jersey
2007 The First Sir William Dunn Scholar, Cambridge
2006 Randolph T. Major Medal: Merck & Company and The University of Connecticut
2002 Frontiers in Biological Chemistry Award: Max Planck Institute Mulheim, Germany
2001 Senior Visiting Fellowship, St. John's College, Oxford
1999 John Morgan Society
1995 IBM Partnership Award
1994 Keilin Medal of the Biochemical Society